Páginas personales


Juan Pablo Gutiérrez García

Catedrático de Mejora Genética Animal


Doctor en Veterinaria


Ingeniero Técnico en Informática de Gestión


Especialista Universitario en Mejora Genética Animal

Departamento de Producción Animal

Facultad de Veterinaria

Universidad Complutense de Madrid

Avda. Puerta de Hierro s/n

28040 Madrid

Phone and FAX: +34 91 394 3767

E-mail: gutgar@vet.ucm.es


pdf FREE!


Endog download area

Molkin download area





Endog 4.8: A computer program for analysing pedigree information


Primary functions carried out by ENDOG are the computation of the individual inbreeding (F) (Wright, 1931) and the average relatedness (AR) (Gutiérrez et al., 2003; Goyache et al., 2003) coefficients. Additionally, users can compute with ENDOG useful parameters in population genetics such as that described for Biochard et al. (1997) for the number of ancestors explaining genetic variability or those proposed by Robertson (1953) and Vassallo et al. (1996) for the genetic importance of the herds. ENDOG also can compute F statistics (Wright, 1978) from genealogical information following Caballero and Toro (2000; 2002). Moreover, the present version of ENDOG calculates effective population size following different methodologies including regression approaches and particularly the recently proposed realized effective population size from individual increase in inbreeding (Gutiérrez et al., 2008), modified to account for avoidance of self-fertilization (Gutiérrez et al., 2009).




Most of them are solved simply by modifying the path of installation by default.


Use R code by Paul Sunnucks (THANKS!)

Or try OpenOffice software

Good luck!


Endog 4.8 User’s Guide (10 november 2010)


Endog 4.8  (English Version) (10 november 2010)


-> Download this additional file for Selfing Version (USER’S GUIDE NO UPDATED!) (28 september 2011)


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Molkin 3.0: A computer program for genetic analysis of populations using molecular coancestry information


MolKin (current version 3.0) is a population genetics computer program that conducts several genetic analyses on multilocus information in a user-friendly environment. Primary functions carried out by MolKin are the computation of the between individuals (and populations) molecular coancestry coefficients (fij), the Kinship distance (Dk) at individual and population levels. Additionally, users can compute with MolKin a set of among populations, genetic distances and F-statistics (Wright, 1978) from multilocus information. The current version of MolKin provides different methods to quantify contributions to diversity to set priorities for conservation. The program will help researchers or those responsible for population management to assess genetic variability and population structure at reduced costs with respect to dataset preparation.


Molkin 3.0 User’s Guide (22 may 2009)


Molkin 3.0 (22 may 2009)


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Para recibir copias escribe un mensaje electrónico

For reprints, please e-mail me


·        ARIAS, K.D., FERNÁNDEZ, I., GUTIÉRREZ, J.P., ÁLVAREZ, I.,  GOYACHE, F., (2024)  Population dynamics of potentially harmful haplotypes: a pedigree analysis. BMC Genomics, 25:487.  [PDF]


·        MOKHTARI,M., ESMAILIZADEH, A., ROUDBARI, Z., BARAZANDEH, A., GUTIÉRREZ, J.P., MOHEBBINEJAD, E., (2024) Early growth performance in the Murciano-Granadina goats: Insights from genetic and phenotypic analyses. Journal of Agricultural Science, in press.  [PDF]


·        CRUZ, A., MURILLO, Y, BURGOS, A., YUCRA, A., MORANTE, R., QUISPE, M., QUISPE, C. QUISPE, E., GUTIÉRREZ, J.P., (2024). Genetic parameters for different types of medullated fiber in alpacas. Journal of Animal Breeding and Geneticsin press. [PDF]


·        MOKHTARI,M., ESMAILIZADEH, A., MIRMAHMOUDI. R., ROUDBARI, Z., BARAZANDEH, A., GUTIÉRREZ, J.P., MOHEBBINEJAD, E., (2024) Genetic and phenotypic analysis of reproductive traits in the Murciano-Granadina does: predictive ability of the statistical models and estimation of genetic parameters. Small Ruminant Research, 232: 107221[PDF]


·        DOMÍNGUEZ, S, C., CERVANTES, I., GUTIÉRREZ, J.P., MORENO, E., (2024) Pedigree analysis in the mhorr gazelle (Nanger dama mhorr): Genetic variability evolution of the captive population. Ecology and Evolution, 14:e10876. [PDF]


·        ARIAS, K.D., HANBOREUM, L, BOZZI, R., ÁLVAREZ, I., GUTIÉRREZ, J.P., FERNANDEZ, I., MENÉNDEZ, J., BEJA-PEREIRA, A., GOYACHE, F., (2024)  Ascertaining the genetic background of the Celtic-Iberian pig strain: a signatures of selection approach. Journal of Animal Breeding and Genetics, 141: 96-112. [PDF]


·        OJEDA-MARÍN, C, GUTIÉRREZ, JP, FORMOSO-RAFFERTY, N, GOYACHE, F, CERVANTES, I, (2024)  Differential patterns in Runs of Homozygosity in two mice lines under divergent selection for environmental variability for birth weight. Journal of Animal Breeding and Genetics, 141: 193-206.[PDF]


·        OJEDA-MARÍN, C., CERVANTES, I., FORMOSO-RAFFERTY, N., GUTIÉRREZ, J.P. (2023) Genomic inbreeding measures applied to a population of mice divergently selected for birth weight environmental variance. Frontiers in  Genetics, 14:1303748.[PDF]


·        MORE, M, VELI, E, CRUZ, A., GUTIÉRREZ, J.P., GUTIÉRREZ, G., PONCE DE LEÓN, A., (2023) Genome Wide Association Study for Fiber Diameter in Alpacas. Animals, 13: 3316.[PDF]


·        EL-OUAZIZI, L., FORMOSO-RAFFERTY, N., CERVANTES, I., GUTIÉRREZ, J.P., (2023) Differential sensitivity of climate conditions on birth weight genetic values in mice divergently selected for birth weight environmental variability. Journal of Animal Science, 101: 1-8.[PDF]


·        ARIAS, K.D., GUTIÉRREZ, J.P., FERNÁNDEZ, I., ÁLVAREZ, I.,  GOYACHE, F., (2023)  Approaching autozygosity in a small pedigree of Gochu Asturcelta pigs. Genetics, Selection and Evolution, 55: 74.  [PDF]


·        MOKHTARI,M., ESMAILIZADEH, A., MIRMAHMOUDI. R., GUTIÉRREZ, J.P., MOHEBBINEJAD, E., (2023) Comparison of non-linear models and genetic parameter estimation for growth curve traits in the Murciano-Granadina goat breed. Small Ruminant Research, 226: 107059. [PDF]


·        ARIAS, K.D., GUTIÉRREZ, J.P., FERNÁNDEZ, I., ÁLVAREZ, I.,  GOYACHE, F., (2023). Copy Number Variation Regions differing in segregation patterns span different sets of genes. Animals, 13, 2351. [PDF]


·        GUTIÉRREZ, J.P., CRUZ, A., MORANTE, B., BURGOS, A., FORMOSO-RAFFERTY, N., CERVANTES, I., (2023). Genetic parameters for fleece uniformity in alpacas. Journal of Animal Science, 101: 1-8.



·        FORMOSO-RAFFERTY, N., EL-OUAZIZI, L., ARIAS-ÁLVAREZ, M., GUTIÉRREZ, J.P., CERVANTES, I., (2023) Embryo survival and fertility differ in lines divergently selected for birth weight homogeneity in mice. Journal of Animal Breeding and Genetics, 00:1-9. [PDF]


·        CRUZ, A., SEDANO, J., BURGOS, A., GUTIÉRREZ, J.P., WURZINGER, M., GUTIÉRREZ-REYNOSO, G., (2023). Genomic selection improves genetic gain for fiber traits in a breeding program for alpacas. Livestock Science, 270: 105195. [PDF]


·        ARIAS, K.D., GUTIÉRREZ, J.P., FERNÁNDEZ, I., MENÉNDEZ-ARIAS, N.A., ÁLVAREZ, I.,  GOYACHE, F., (2023)  Segregation patterns and inheritance rate of Copy Number Variations regions assessed in a Gochu Asturcelta pig pedigree. Gene, 854: 14711.[PDF] 


·        ARIAS, K.D., ÁLVAREZ, I., GUTIÉRREZ, J.P., FERNÁNDEZ, I., MENÉNDEZ, J., MENÉNDEZ-ARIAS, N.A., GOYACHE, F., (2022)  Understanding Mendelian Errors in SNP arrays data using a Gochu Asturcelta pig pedigree: genomic alterations, family size and calling errors. Scientific Reports, 12: 19686. [PDF]


·        FORMOSO-RAFFERTY, N., GUTIÉRREZ, J.P., GARCÍA-ÁLVAREZ, A, PÉREZ, T., CERVANTES, I., , (2022)  Impact of selection for birth weight variability on reproductive longevity: a mice model. Journal of Animal Breeding and Genetics, 139:370–379. [PDF]


·        MANCISIDOR, B., CRUZ, A., GUTIÉRREZ, G.A., BURGOS, A., MORÓN, J., WURZINGER, M., GUTIÉRREZ, J.P., (2021) ss-GBLUP method improves the accuracy of breeding value prediction in Huacaya alpaca. Animals, 11: 3052. [PDF]


·        ARIAS, K.D., CERVANTES, I., GUTIÉRREZ, J.P., (2021) Impact of the event effect in genetic evaluation for ranking traits in horses. Journal of Animal Breeding and Genetics, 139: 13-25. [PDF]


·        CRUZ, A., YUCRA, A., GUTIÉRREZ, G.A., BURGOS, A., MORANTE, R., GUTIÉRREZ, J.P., CERVANTES, I., WURZINGER, M., (2021) Colorimetry analysis of coat color and its relationship with fiber traits in alpacas. Animal, 15(5): 100219. [PDF]


·        CRUZ, A., GUTIÉRREZ, G.A., BURGOS, A., MORANTE, R., WURZINGER, M., GUTIÉRREZ, J.P., (2021) Management software for pedigree and production of New World Camelids: Pacokipu and Llamakipu. Revista de Investigaciones Veterinarias del Perú, 32(4): e19355. [PDF]


·        OJEDA-MARÍN, C., CERVANTES, I., MORENO, E., GOYACHE, F., GUTIÉRREZ, J.P., (2021) Breeding strategies to optimise effective population size in low census captive populations: the Gazella cuvieri case. Animals, 11, 1559. [PDF]


·        GARCÍA-BALLESTEROS, S., VILLANUEVA, B., FERNÁNDEZ, J., GUTIÉRREZ, J.P., CERVANTES, I., (2021) Genetic parameters for uniformity of harvest weight in Pacific white shrimp (Litopenaeus vannamei). Genetics, Selection and Evolution, 53:26. [PDF]


·        PINARES, R., CRUZ, A., DAVERIO, M.A., GUTIÉRREZ, J.P., PONCE DE LEÓN, F.A., WURZINGER, M., DI ROCCO, F., GUTIÉRREZ, G.A., 2021. Single nucleotide polymorphisms (SNPs) of the MC1R gene in black and brown alpacas. Revista peruana de biología, 28: e19742. [PDF]


·        POYATO-BONILLA, J., SÁNCHEZ-GUERRERO, M.J., CERVANTES, I., GUTIÉRREZ, J.P., VALERA, M., (2021) Genetic parameters for canalisation analysis of morphological traits in the Pura Raza Español horse. Journal of Animal Breeding and Genetics, 138: 482– 490. [PDF]


·        CERVANTES, I., GUTIÉRREZ, J.P., GARCÍA-BALLESTEROS, S., VARONA, L., (2020) Combining threshold, thurstonian and classical linear models in horse genetic evaluations for endurance competitions. Animals, 10: 1075. [PDF]


·        DOMÍNGUEZ-VIVEROS, J., RODRÍGUEZ-ALMEIDA, F.A., MEDELLÍN-CÁZARES, A., GUTIÉRREZ, J.P., (2020). Pedigree analysis in ten sheep populations in Mexico. Revista Mexicana de Ciencias Pecuarias, 11: 1071-1086. [PDF]


·        CRUZ, A., GUTIÉRREZ, J.P., TORRES, R., FORMOSO-RAFFERTY, N., MORANTE, R., BURGOS, A., CERVANTES, I., (2020) Calving date and its variability as a potential trait in the breeding objective to account for reproductive seasonality in alpacas. Reproduction in Domestic Animals, 55: 814-821. [PDF]


·        FORMOSO-RAFFERTY, N., CHÁVEZ, K.N., OJEDA, C., CERVANTES, I., GUTIÉRREZ, J.P., (2020)  Selection response in a divergent selection experiment for birth weight variability in mice compared with a control line. Animals, 10: 920. [PDF]


·        CERVANTES, I, BODIN, L., VALERA, M., MOLINA, A., GUTIÉRREZ, J.P., (2020) Challenging the selection for consistency in the rank of endurance competitions. Genetics, Selection and Evolution, 52:20. [PDF]


·        CRUZ, A., MENÉNDEZ-BUXADERA, A., GUTIÉRREZ, G., MORANTE, R., BURGOS, A., GUTIÉRREZ, J.P., (2020) Genetic (co) variance across age of fiber diameter and standard deviation in Huacaya alpacas, estimated by repeatability, multi-trait and random regression models. Livestock Science, 23, UNSO 103863. [PDF]


·        TATLIYER, A., CERVANTES, I., FORMOSO-RAFFERTY, N., GUTIÉRREZ, J.P., (2019)  The Statistical Scale Effect as a Source of Positive Genetic Correlation between Mean and Variability: A Simulation Study. G3: Genes Genomes Genetics, 9: 3001-3008. [PDF]


·        FORMOSO-RAFFERTY, N., CERVANTES, I., SÁNCHEZ, J.P., GUTIÉRREZ, J.P., BODIN, L., (2019)  Effect of feed restriction on the environmental variability of birth weight in divergently selected lines of mice. Genetics, Selection and Evolution, 2019, 51:27. [PDF]


·        MOKHTARI, M.S., ASADI FOZI, M., GUTIERREZ, J.P., NOTTER, D.R., (2019) Genetic and phenotypic aspects of early reproductive performance in Raeini Cashmere goat. Tropical Animal Health and Production, 51: 2175-2180. [PDF]


·        CRUZ, A., MORANTE, R., GUTIÉRREZ, J.P., TORRES, R., BURGOS, A., CERVANTES, I., (2019) Genetic parameters for medullated fiber and its relationship with other productive traits in alpacas. Animal, 13: 1358-1364. [PDF]


·        PINARES, R., GUTIÉRREZ, G.A., CRUZ, A., BURGOS, A., GUTIÉRREZ, J.P., (2019) Phenotypic variability of percentage of medullated fibers in Huacaya alpaca fleece. Revista de Investigaciones Veterinarias del Perú, 30: 699-708. [PDF]


·        FORMOSO-RAFFERTY, N., DE LA FLOR, M., GUTIÉRREZ, J.P., CERVANTES, I., (2018)  Feed and reproductive efficiency differences between divergently selected lines for birth weight environmental variability in mice. Journal of Animal Breeding and Genetics, 135: 378-389. [PDF]


·        PINARES, R., GUTIÉRREZ, G.A., CRUZ, A., MORANTE, R., CERVANTES, I., BURGOS, A., GUTIÉRREZ, J.P., (2018) Heritability of individual fiber medullation in Peruvian alpacas. Small Ruminant Research,  165: 93-100. [PDF]


·        GARCÍA-BALLESTEROS, S., VARONA, L., VALERA, M., GUTIÉRREZ, J.P., CERVANTES, I., (2018) Cross-validation analysis for genetic evaluation models for ranking in endurance horses. Animal, 12: 20-27. [PDF]


·        GARCÍA-BALLESTEROS, S., GUTIÉRREZ, J.P., VARONA, L., FERNÁNDEZ, J., (2017) The influence of natural selection in breeding programs: a simulation study.. Livestock Science, 204: 98-103. [PDF]


·        SÁNCHEZ-GUERRERO, M.J., CERVANTES, I., MOLINA, A., GUTIÉRREZ, J.P., VALERA, M., (2017) Designing an early selection morphological linear traits index for dressage in the Pura Raza Español horse. Animal, 11: 948-957. [PDF]


·        CRUZ, A., CERVANTES, I., BURGOS, A., MORANTE, R., GUTIÉRREZ, J.P., (2017) Genetic parameters estimation for preweaning traits and their relationship with reproductive, productive and morphological traits in alpaca. Animal, 11: 746-754. [PDF]


·        FORMOSO-RAFFERTY, N., CERVANTES, I., IBÁÑEZ-ESCRICHE, N., GUTIÉRREZ, J.P., (2017) Modulating birth weight heritability in mice. Journal of Animal Science, 95:531-537. [PDF]


·        CRUZ, A. , MORANTE, R., CERVANTES, I., BURGOS, A., GUTIÉRREZ, J.P., (2017) Effect of the gestation and lactation on fiber diameter and its variability in Peruvian alpacas. Livestock Science, 198: 31-36. [PDF]


·        FORMOSO-RAFFERTY, N., CERVANTES, I., IBÁÑEZ-ESCRICHE, N., GUTIÉRREZ, J.P., (2016) Correlated genetic trends for production and welfare traits in a mouse population divergently selected for birth weight environmental variability. Animal, 10(11): 1770-1777. [PDF]


·        CERVANTES, I., GUTIÉRREZ, J.P., MEUWISSEN, T.H.E., (2016) Response to selection while maximizing genetic variance in small populations. Genetics, Selection and Evolution, 48: 69. [PDF]


·        CERVANTES. I., BARTOLOMÉ, E., VALERA, M., GUTIÉRREZ, J.P., MOLINA, A., (2016) Crossbreed genetic performance study in the eventing horse competition. Animal Production Science, 56: 1454-1462. [PDF]


·        FORMOSO-RAFFERTY, N., CERVANTES, I., IBÁÑEZ-ESCRICHE, N., GUTIÉRREZ, J.P., (2016) Genetic control of the environmental variance for birth weight in seven generations of a divergent selection experiment in mice. Journal of Animal Breeding and Genetics, 133: 227-237. [PDF]


·        MARTÍN DE LA ROSA, A.J., CERVANTES, I., GUTIÉRREZ, J.P., (2016) The equivalent effective population size mating as a useful tool in the genetic management of the ibicenco rabbit breed (conill pages d'eivissa). Czech Journal of Animal Science, 61: 108-116. [PDF]


·        MAMANI-CATO, R., ACERO, R.G., MAMANI, T.H., GUTIÉRREZ, J.P., (2016) Genetic structure of the population of llamas (Lama glama) of Gene bank National Institute of Innovation Agrarian-Peru. Journal of High Andean Research, 18: 55-60. [PDF]


·        CRUZ, A., CERVANTES, I., BURGOS, A., MORANTE, R., GUTIÉRREZ, J.P., (2015) Estimation of genetic parameters for reproductive traits in alpacas. Animal Reproduction Science, 163: 48-55. [PDF]


·        MOKHTARI, M.S., MIRAEI-ASHTIANI, S. R., JAFAROGHLI, M., GUTIÉRREZ, J.P., (2015) Studying the Genetic Diversity in Moghani Sheep Using Pedigree Analysis. Journal of Agricultural Science and Technology, 17: 1151-1160. [PDF]


·        CANAZA-CAYO, A.W., HUANCA, T., GUTIÉRREZ, J.P., BELTRÁN, P.A., (2015) Modelling of growth curves and estimation of genetic parameters for growth curve parameters in Peruvian young llamas (Lama glama). Small Ruminant Research, 130: 81–89. [PDF]


·        RASHIDI A, MOKHTARI, M.S., GUTIÉRREZ, J.P., (2015) Pedigree analysis and inbreeding effects on early growth traits and greasy fleece weight in Markhoz goat. Small Ruminant Research, 124: 1-8. [PDF]


·        IBÁÑEZ, B., CERVANTES, I., GUTIÉRREZ, J.P., GOYACHE, F., MORENO, E., (2014) Estimates of direct and indirect effects for early juvenile survival in captive populations maintained for conservation purposes: the case of Cuvier's gazelle. Ecology and Evolution, 4(21): 4117–4129. [PDF]


·        DELGADO, J.F., DE ANDRÉS, N., VALERA, M, GUTIÉRREZ, J.P., CERVANTES, I., (2014) Assessment of population structure depending on breeding objectives in Spanish Arabian horse by genealogical and molecular information. Livestock Science, 168: 9-16. [PDF]


·        SÁNCHEZ, M.J., CERVANTES, I., VALERA, M., GUTIÉRREZ, J.P., (2014) Modelling genetic evaluation for dressage in Pura Raza Español horses with focus on the rider effect. Journal of Animal Breeding and Genetics, 131: 395-402. [PDF]


·        SOLÉ, M., CERVANTES, I., GUTIÉRREZ, J.P., GÓMEZ, M.D., VALERA, M., (2014) Estimation of genetic parameters for morphological and functional traits in a Menorca horse population. Spanish Journal of Agricultural Research, 12: 125-132. [PDF]


·        GUTIÉRREZ, J.P., CERVANTES, I., PÉREZ-CABAL, M.A., BURGOS, A., MORANTE, R., (2014) Weighting fibre and morphological traits in a genetic index for an alpaca breeding program. Animal, 8(3): 360-369. [PDF]


·        PAREDES, M.M., MEMBRILLO, A., GUTIÉRREZ, J.P., CERVANTES, I., AZOR, P.J., MORANTE, R., ALONSO, A., MOLINA, A., MUÑOZ-SERRANO, A., (2014) Association of microsatellite markers with fiber diameter trait in Peruvian alpacas (Vicugna pacos). Livestock Science, 161: 6-16. [PDF]


·        MOKHTARI M.S., MORADI SHAHRBABAK M., ESMAILIZADEH, A. K., MORADI SHAHRBABAK, H., GUTIÉRREZ, J.P., (2014) Pedigree analysis of Iran-Black sheep and inbreeding effects 1 on growth and reproduction traits. Small Ruminant Research, 116: 14– 20. [PDF]


·        PÉREZ-CABAL, M.A., LEGAZ, E., CERVANTES, I., DE LA FUENTE, L. F., MARTÍNEZ, R., GOYACHE, F., GUTIÉRREZ, J.P., (2013) Association between body and udder morphological traits and dairy performance in Spanish Assaf sheep. Archives of Animal Breeding, 56:430-442. [PDF]


·        MOKHTARI M.S., MORADI SHAHRBABAK M., ESMAILIZADEH, A. K., ABDOLLAHI-ARPANAHI, R., GUTIÉRREZ, J.P., (2013) Genetic diversity in Kermani sheep assessed from pedigree analysis. Small Ruminant Research, 114: 202-205. [PDF]


·        PUN, A., CERVANTES, I., NIETO, B., SALGADO, C., PÉREZ-CABAL, M.A., IBÁÑEZ-ESCRICHE, N., GUTIÉRREZ, J.P., (2013) Genetic parameters for birth weight environmental variability in mice. Journal of Animal Breeding and Genetics, 130: 404-414. [PDF]


·        KORRIDA, A., GUTIÉRREZ, J.P., AGGREY, S.E., AMIN-ALAMI, A., (2013) Genetic variability characterization of the Moroccan Houbara Bustard (Chlamydotis undulata undulata) inferred from pedigree analysis. Zoo Biology, 32: 366-373. [PDF]


·        MORENO, A., IBÁÑEZ-ESCRICHE, N., GARCÍA-BALLESTEROS, S., SALGADO, C., NIETO, B., GUTIÉRREZ, J.P., (2012) Correlated genetic trend in the environmental variability of weight traits in mice. Livestock Science, 148: 189–195. [PDF]


·        SANTANA JR, M.L., OLIVEIRA, P.S., ELER, J.P., FERRAZ, J.B.S., GUTIÉRREZ, J.P., (2012) Pedigree analysis and inbreeding depression on growth traits in Brazilian Marchigiana and Bonsmara breeds. Journal of Animal Science, 90: 99-108. [PDF]


·        PUN, A., GOYACHE, F., CERVANTES, I., GUTIÉRREZ, J.P., (2012) Cytoplasmic line effects for birth weight and preweaning growth traits in the Asturiana de los Valles beef cattle breed. Livestock Science,143: 177-183. [PDF]


·        BARTOLOMÉ, E., CERVANTES, I., VALERA, M., GUTIÉRREZ, J.P., (2011) Influence of foreign breeds on the genetic structure of the Spanish Sport Horse population. Livestock Science, 142: 70-79. [PDF]


·        LEGAZ, E., CERVANTES, I., PÉREZ-CABAL, M.A., DE LA FUENTE, L.F., MARTÍNEZ, R., GOYACHE, F., GUTIÉRREZ, J.P., (2011) Multivariate characterisation of morphological traits in Assaf (Assaf.E) sheep. Small Ruminant Research, 100: 122-130. [PDF]


·        MORENO, A., SALGADO, C., PIQUERAS, P., GUTIÉRREZ, J.P., TORO, M.A., IBÁÑEZ-ESCRICHE, N., NIETO, B., (2011) Restricting inbreeding while maintaining selection response for weight gain in Mus musculus. Journal of Animal Breeding and Genetics, 128: 276-283. [PDF]


·        VARONA, L., CERVANTES, I., PÉREZ-CABAL, M.A., MORANTE, R., BURGOS, A., GUTIÉRREZ, J.P., (2011) Genetic parameters for growth of fiber diameter in alpacas. Fibre Production in South American Camelids and Other Fibre Animals. 107-110. [PDF]


·        MORANTE, R., BURGOS, A., GUTIÉRREZ, J.P., (2011) Producing alpaca fibre for the textile industry. Fibre Production in South American Camelids and Other Fibre Animals. 107-110. [PDF]


·        GUTIÉRREZ, J.P., VARONA, L., PUN, A., MORANTE, R., BURGOS, A., CERVANTES, I., PÉREZ-CABAL, M.A., (2011) Genetic parameters for growth of fiber diameter in alpacas. Journal of Animal Science, 89: 2310–2315. [PDF]


·        CERVANTES, I., PASTOR, J.M., GUTIÉRREZ, J.P., GOYACHE, F., MOLINA, A., (2011) Computing Effective Population Size from molecular data: The case of three rare Spanish ruminant populations. Livestock Science, 138:202-206. [PDF]


·        CERVANTES, I., GOYACHE, F., MOLINA, A., VALERA, M., GUTIÉRREZ, J.P., (2011) Estimation of effective population size from the rate of coancestry in pedigreed Populations. Journal of Animal Breeding and Genetics,128: 56-63. [PDF]


·         PÉREZ-CABAL, M.A., CERVANTES, I., MORANTE, R., BURGOS, A., GOYACHE, F., GUTIÉRREZ, J.P., (2010) Analysis of the existence of major genes affecting alpaca fiber traits. Journal of Animal Science, 88: 3783-3788. [PDF]


·        PANETTO, J.C.C., GUTIÉRREZ, J.P., FERRAZ, J.B.S., CUNHA, D.G., GOLDEN, B.L., (2010) Assessment of inbreeding depression in a Guzerat dairy herd: effects of individual increase in inbreeding coefficients on production and reproduction. Journal of Dairy Science, 93: 4902-4912. [PDF]


·        PÉREZ-PARDAL, L., ROYO, L.J., BEJA-PEREIRA, A., CHEN, S., CANTET, R:J.C., TRAORÉ, A., CURIK, I., SÖLKNER, J., BOZZI, R., FERNÁNDEZ, I., ÁLVAREZ, I., GUTIÉRREZ, J.P., GÓMEZ, E., PONCE DE LEON, F.A., GOYACHE, F. (2010) Multiple paternal origins of domestic cattle revealed by Y-specific interspersed multilocus microsatellites. Heredity, 105: 511-519. [PDF] [Supplementary information]


·        BARTOLOMÉ, E., GOYACHE, F., MOLINA, A., CERVANTES. I., VALERA, M. GUTIÉRREZ, J.P. (2010) Pedigree estimation of the (sub) population contribution to the total gene diversity: the horse coat colour case. Animal, 4(6):867-875. [PDF]


·        CERVANTES, I., PÉREZ-CABAL, M.A., MORANTE, R., BURGOS, A., SALGADO, C., NIETO, B., GOYACHE, F., GUTIÉRREZ, J.P. (2010) Genetic parameters and relationships between fibre and type traits in two breeds of Peruvian alpacas. Small Ruminant Research, 88:6-11. [PDF]


·        CERVANTES, I., GUTIÉRREZ, J..P., FERNÁNDEZ, I., GOYACHE, F., (2010) Genetic relationships among calving ease, gestation length, and calf survival to weaning in the Asturiana de los Valles beef cattle breed. Journal of Animal Science, 88: 96-101. [PDF]


·        GOYACHE, F., FERNÁNDEZ, I., ESPINOSA, M.A., PAYERAS, L., PÉREZ-PARDAL, L., GUTIÉRREZ, J.P., ROYO, L.J., ÁLVAREZ, I. (2010) Demographic and genetic analysis of the Mallorquina sheep flockbook. ITEA, 106, 3-14. [PDF]


·        ÁLVAREZ, I., GUTIÉRREZ, J..P., ROYO, L. J., FERNÁNDEZ, I., GOYACHE, F., (2009) Quantifying diversity losses due to selection for scrapie resistance in three endangered Spanish sheep breeds using microsatellite information. Preventive Veterinary Medicine, 91: 172-178. [PDF]


·        CERVANTES, I., GUTIÉRREZ, J.P., MOLINA, A., GOYACHE, F., VALERA, M., (2009) Genealogical analyses in open populations: the case of three Arab-derived Spanish horse breeds. Journal of Animal Breeding and Genetics, 126: 335-347. [PDF]


·        GUTIÉRREZ, J.P., CERVANTES, I., GOYACHE, F., (2009) Improving the estimation of realised effective population sizes in farm animals. Journal of Animal Breeding and Genetics,126: 327-332. [PDF]


·        CERVANTES, I., BAUMUNG, R., MOLINA, A., DRUML, T., GUTIÉRREZ, J.P., SÖELKNER, J., VALERA, M., (2009) Size and shape analysis of morphofunctional traits in the spanish arab horse. Livestock Science, 125: 43-49. [PDF]


·        TRAORE, A., ÁLVAREZ, I., TAMBOURA, H.H., ROYO, L.J., KABORE, A., FERNANDEZ, I., GUTIÉRREZ, J.P., SANGARÉ, M., OUEDRAOGO-SANOU, G., TOGUYENI, A., SAWADOGO, L., GOYACHE, F., (2009) Genetic characterisation of Burkina Faso goats using microsatellite polymorphism. Livestock Science, 123: 322-328. [PDF]


·        GUTIÉRREZ, J.P., GOYACHE, F., BURGOS, A., CERVANTES, I., (2009) Genetic analysis of six production traits in Peruvian alpacas. Livestock Science, 123: 193-197. [PDF]


·        GÓMEZ, M.D., VALERA, M., MOLINA, A.., GUTIÉRREZ, J.P., GOYACHE, F., (2009) Assessment of inbreeding depression for body measurements in Spanish Purebred (Andalusian) horses. Livestock Science, 122: 149-155. [PDF]


·        LEGAZ, E., ÁLVAREZ, I., ROYO, L.J., FERNÁNDEZ, I., GUTIÉRREZ, J.P., GOYACHE, F., (2008) Genetic relationships between Spanish Assaf (Assaf.E) and Spanish native dairy sheep breeds. Small Ruminant Research, 80: 39-44. [PDF]


·        CERVANTES, I., GOYACHE, F., MOLINA, A., VALERA, M., GUTIÉRREZ, J.P., (2008) Application of individual increase in inbreeding to estimate realised effective sizes from real pedigrees. Journal of Animal Breeding and Genetics, 125: 301-310. [PDF]


·        GUTIÉRREZ, J.P., CERVANTES, I., MOLINA, VALERA, M., A., GOYACHE, F., (2008) Individual increase in inbreeding allows estimating effective sizes from pedigrees. Genetics, Selection and Evolution, 40: 359-378. [PDF] The original publication is available at http://www.gse-journal.org


·        IBÁÑEZ-ESCRICHE, N., MORENO, A., NIETO, B., PIQUERAS, P., SALGADO, C., GUTIÉRREZ, J.P., (2008) Genetic parameters related to environmental variability of weight traits in a selection experiment for weight gain in mice; signs of correlated canalised response. Genetics, Selection and Evolution, 40: 279-293. [PDF] The original publication is available at http://www.gse-journal.org


·        ÁLVAREZ, I., ROYO, L.J., GUTIÉRREZ, J.P., FERNÁNDEZ, I., ARRANZ, J.,J., GOYACHE, F., (2008) Relationship between genealogical and microsatellite information characterising losses of genetic variability: empirical evidence from the rare Xalda sheep breed. Livestock Science,115: 80-88. [PDF]


·        DÍAZ-SAN SEGUNDO, F., SEVILLA, N., GUTIÉRREZ, J.P., BRUN, A., (2008) Altered lymphocyte homeostasis after oral prion infection in mouse. Veterinary Inmunology and Immunopathology, 122: 204-215. [PDF]


·        CERVANTES, I., MOLINA, A., GOYACHE, F., GUTIÉRREZ, J.P., VALERA, M., (2008) Population history and genetic variability in the Spanish Arab Horse assessed via pedigree analysis. Livestock Science, 113: 24-33. [PDF]


·        TRAORÉ, A., ROYO, L.J., ÁLVAREZ, I., FERNÁNDEZ, I, GUTIÉRREZ,  J.P.,  RINCÓN, C., PÉREZ-PARDAL, L., GOYACHE, F., (2008). Adjacent mutations to ovine PrnP codons 136 and 171 affect performance of the diagnostic protocol based on RT-PCR coupled to fluorescence probes. ITEA, 104(2), 106-109. [PDF]


·        ÁLVAREZ, I., ROYO, L.J., FERNÁNDEZ, I., GUTIÉRREZ,  J.P., PÉREZ-PARDAL, L., GUERRA, V., RINCÓN, C., TRAORÉ, A., GOYACHE, F., (2008). Genetic differentiation between two geographic subpopulations of Bermeya goat. ITEA, 104(2), 290-294. [PDF]


·        CERVANTES, I., GOYACHE, F., GUTIÉRREZ,  J.P.,  (2008). Ratio between inbreeding and coancestry rates as a measure of population subdivision. Preliminary results. ITEA, 104(2), 303-307. [PDF]


·        AZOR, P.J., CERVANTES, I., VALERA, M., ARRANZ, J.J., MEDINA, C., GUTIÉRREZ,  J.P., GOYACHE, F., MUÑOZ, A., MOLINA, A., (2008). Preliminary assessment of population structure of Spanish Merino Breed: traditional strains situation using genealogical and molecular analysis. ITEA, 104(2), 295-302. [PDF]


·        GONZÁLEZ-RECIO, O., LÓPEZ DE MATURANA, E., GUTIÉRREZ, J.P., (2007) Inbreeding depression on female fertility and calving ease in Spanish dairy cattle. Journal of Dairy Science, 90: 5744-5752. [PDF]


·        ÁLVAREZ,  I., ROYO, L.J., GUTIÉRREZ, J.P., FERNÁNDEZ, I., ARRANZ, J.,J., GOYACHE, F., (2007) Genetic diversity loss due to selection for scrapie resistance in the rare Spanish Xalda sheep breed. Livestock Science, 111: 204-212. [PDF]


·        GUTIÉRREZ, J.P., LEGAZ, E., GOYACHE, F., (2007) Genetic parameters affecting 180-days standardised milk yield, test-day milk yield and lactation length in Spanish Assaf (Assaf.E) dairy sheep.  Small Ruminant Research, 70: 233-238. [PDF]


·        ROYO, L.J., ÁLVAREZ,  I., GUTIÉRREZ, J.P., FERNÁNDEZ, I., GOYACHE, F., (2007) Genetic variability in the endangered Asturcón pony assessed using genealogical and molecular information. Livestock Science, 107: 162-169.  [PDF]


·        GUTIÉRREZ, J.P., GOYACHE, F., FERNÁNDEZ, I.,  ÁLVAREZ,  I., ROYO, L.J., (2007) Genetic relationships among calving ease, calving interval, birth weight and weaning weight in the Asturiana de los Valles beef cattle breed. Journal of Animal Science, 85: 69-75. [PDF]


·        GUTIÉRREZ, J.P., NIETO, B., PIQUERAS, P., IBÁÑEZ, N., SALGADO, C., (2006) Genetic parameters for canalisation analysis of litter size and litter weight traits at birth in mice. Genetics, Selection and Evolution, 38: 445-462. [PDF]


·        GUTIÉRREZ, J.P., FERNÁNDEZ, I., ALVAREZ, I., ROYO, L.J., GOYACHE, F., (2006) Sire x contemporary group interactions for birth weight and preweaning growth traits in the Asturiana de los Valles beef cattle breed.  Livestock Science, 99: 61-68. [PDF]


·        GUTIÉRREZ, J.P., ROYO L.J., ÁLVAREZ, I., GOYACHE, F., (2005) Molkin v2.0: a computer program for genetic analysis of populations using molecular coancestry information. Journal of Heredity, 96: 718-721. [PDF]


·        ROYO L.J., ÁLVAREZ, I., BEJA-PEREIRA, A., MOLINA, A., FERNÁNDEZ, I., JORDANA, J., GÓMEZ, E., GUTIÉRREZ, J.P., GOYACHE, F., (2005). The origins of Iberian horses assessed via mitochondrial DNA. Journal of Heredity, 96: 663-669. [PDF]


·        GUTIÉRREZ, J.P., MARMI, J., GOYACHE, F., JORDANA, J., (2005) Pedigree information reveals moderate to high levels of inbreeding and a population genetic structure in the Catalonian donkey breed. Journal of Animal Breeding and Genetics, 122: 378-386. [PDF]


·        GUTIÉRREZ, J.P., GOYACHE, F. (2005) A note on ENDOG: a computer program for analysing pedigree information. Journal of Animal Breeding and Genetics, 122: 172-176. [PDF]


·        ALVAREZ I., GUTIÉRREZ, J.P., ROYO, L.J., FERNÁNDEZ, I., GOMEZ,E.,  ARRANZ, J.,J., GOYACHE, F., (2005) Testing the usefulness of the molecular coancestry information to assess genetic relationships on livestock using a set of Spanish sheep breeds. Journal of Animal Science, 83: 737-744. [PDF]


·        VALERA, M., MOLINA, A., GUTIÉRREZ, J.P., GÓMEZ, J.,  GOYACHE, F. (2005) Pedigree analysis in the Andalusian horse: population structure, genetic variability and influence of the Carthusian strain. Livestock Production Science, 95: 57-66. [PDF]


·        GOYACHE, F., GUTIÉRREZ, J.P., FERNÁNDEZ, I.,  ROYO, L.J., ÁLVAREZ,  I., GÓMEZ, E. (2005) Genetic analysis of days open in Asturiana de los Valles beef cattle breed. Livestock Production Science, 93: 283-289. [PDF]


·        ALVAREZ I., ROYO, L.J., FERNÁNDEZ, I., GUTIÉRREZ, J.P., GOMEZ,E.,  GOYACHE, F. (2004) Genetic relationships and admixture among sheep breeds from Northern Spain assessed using microsatellites. Journal of Animal Science, 82, 2246-2252.[PDF]


·        GOYACHE, F., GUTIÉRREZ, J.P., ÁLVAREZ I., FERNÁNDEZ I., ROYO L.J, GOMEZ, E. (2003) Genetic analysis of calf survival at different preweaning ages in beef cattle. Livestock Production Science, 83: 13–20. [PDF]


·        GOYACHE F.; FERNÁNDEZ I., ROYO L.J, ÁLVAREZ I. GUTIÉRREZ, J.P. (2003) Factors affecting actual weaning weight, preweaning average daily gain and relative growth rate in Asturiana de los Valles beef cattle breed. Archives of Animal Breeding, 46: 235-244. [PDF]


·        GUTIÉRREZ, J.P., ALTARRIBA, J., DÍAZ, C., QUINTANILLA, R., CAÑÓN, J. PIEDRAFITA, J. (2003) Pedigree analysis of eight Spanish beef cattle breeds. Genetics, Selection and Evolution, 35: 43-63. [PDF]


·        GOYACHE, F., GUTIÉRREZ, J.P., FERNANDEZ, I., GOMEZ, E.,  ALVAREZ, I. DIEZ, J.  ROYO, L.J. (2003) Using pedigree information to monitor genetic variability of endangered populations: the Xalda sheep breed of Asturias as an example. Journal of Animal Breeding and Genetics, 120: 95-103. [PDF]


·        GUTIÉRREZ, J.P., ALVAREZ, I, FERNÁNDEZ, I, ROYO, L. J, DÍEZ, J., GOYACHE, F. (2002) Genetic relationships between calving date, calving interval, age at first calving and type traits in beef cattle. Livestock Production Science, 78: 215-222. [PDF]


·        GUTIÉRREZ, J.P., GOYACHE, F. (2002) Estimation of genetic parameters of type traits in Asturiana de los Valles beef cattle breed. Journal of Animal Breeding and Genetics, 119:  93-129. [PDF]


·        GOYACHE, F., ROYO, L.J., ALVAREZ, I., GUTIÉRREZ, J.P. (2002) Testing a continuous variation in preweaning expression of muscular hypertrophy in beef cattle using field data. Archives of Animal Breeding, 45 (2): 139-149. [PDF]


·        GOYACHE, F., GUTIÉRREZ, J.P. (2001) Heritabilities of reproductive traits in Asturiana de los Valles beef cattle breed. Archives of Animal Breeding, 44 (5): 489-496. [PDF]


·        FERNANDEZ, J., MORENO, A., GUTIÉRREZ, J.P., NIETO, B., PIQUERAS, P., SALGADO, C. (1998) Direct and correlated response for litter size and litter weight at birth in the first parity in mice. Livestock Production Science. 53: 217-223. [PDF]


·        DUNNER S., CHECA M.L.., GUTIÉRREZ J.P., MARTÍN J.P., CAÑÓN J. (1998) Genetic analysis and management in small populations: the Asturcon pony as an example, Genetics, Selection and Evolution, 30: 397-405. [PDF]


·        GUTIÉRREZ, J.P., CAÑÓN, J., GOYACHE, F. (1997) Estimation of direct and maternal genetic parameters for preweaning traits in the Asturiana de los Valles beef cattle breed through animal and sire models. Journal of Animal Breeding and Genetics, 114, 261-266. [PDF]


·        GUTIÉRREZ, J.P., CAÑON, J., RICO, M. (1994) Comparison of two models for estimation of variance components in a sample of Spanish Holstein Friesians. Journal of Animal Breeding and Genetics, 111: 169-174. [PDF]


·        CAÑÓN J., GUTIÉRREZ, J.P., DUNNER S., GOYACHE F., VALLEJO M. (1994) Herdbook analysis of the Asturiana beef cattle breeds. Genetics, Selection and Evolution, 26: 65-75. [PDF]


·        CAÑON, J., GRUAND, J., GUTIÉRREZ, J.P., OLLIVIER, L. (1992) Expérience de sélection sur la croissance du tissu maigre chez le porc, avec un dispositif de pères répétés: evolution génétique des caractères soumis à la sélection. Genetics, Selection and Evolution, 24: 449-462. [PDF]



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Félix Goyache